The 49ers pieced together an uneven performance in their preseason dress rehearsal, with dropped passes and penalties looming large in their 23-17 defeat Saturday against the Indianapolis Colts.

Jimmy Garoppolo played the entire first half and the first possession of the third quarter, but failed to orchestrate a touchdown drive. San Francisco instead settled for Robbie Gould field goals when passes fell to the turf after clanging off the hands of Garoppolo’s intended targets.

Here are five takeaways from Saturday’s defeat.

49ers running game gets going

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