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49ers sign linebacker Wheeler, release three

SANTA CLARA -- The 49ers on Thursday signed veteran inside linebacker Philip Wheeler to a one-year contract, and released three players from their 90-man roster.

The 49ers waived wide receiver Trindon Holliday, guard Fouimalo Fonoti and wide receiver Lance Lewis.

Wheeler, 30, a seven-year veteran, does not count against the formula for compensatory picks next year because the Miami Dolphins released him this offseason.

The Dolphins signed Wheeler to a five-year, $26 million contract in 2013, and paid him $13 million in the first two years of that deal. But the seven-year veteran started just four of the 15 games in which he appeared last season.