A lot is on the line when Iowa travels to Happy Valley at 3:30 p.m. Saturday to take on Penn State: The Hawkeyes are still trying to win the West, while the Nittany Lions are hoping to win out and earn a spot in the Rose Bowl.

Here are the key matchups to Saturday’s game that we think should go a long way in determining the winner:

John McGonigal: Penn State’s first-down run defense vs. Iowa

Brent Pry’s unit has a first-down problem, and Iowa will try to take advantage.

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The Nittany Lions’ defense has not played the run particularly well.