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Why did Bolts run D flop? Bosa missed?

In their exhibition opener Saturday, Chargers backup defenders got stampeded.

The Tennessee Titans finished with 288 yards rushing off 8.7 per carry.

The starters handled the run aside from Demarco Murray's 15-yard gain off a counter rush.

Several second- and third-string defenders struggled to get off blocks, take good ankles or make open-field tackles.

The performance gets a yellow flag -- not a bright red one -- because of various circumstances.

The second unit was without defensive ends Tenny Palepoi and Sean Lissemore, who had missed the past five and four practices, respectively. Also unavailable was rookie Jatavis Brown, a speedy linebacker.